Solanaceae Source

A global taxonomic resource for the nightshade family

Solanum remyanum

Citation author: 
Phil.
Citation: 
Anal. Univ. Chil. 91: 17. 1895.
Type: 
Based on Witheringia flexuosa Remy.
Last edited by: 
Bennett, J.
Written by: 
Bennett, J.
Habit: 
Woody shrub 20–100 cm high. Young stems green, older stems brown, woody, subglabrous to densely pubescent, the hairs erect, simple, uniseriate, unicellular or occasionally multicellular, up to 0.2 mm long.
Sympodial structure: 
Sympodial units plurifoliate, not geminate.
Leaves: 
Leaves pinnate or bipinnate, fleshy, the blades 3.3–8.5 x 0.8–5.5 cm, 0.95–2.0 times as long as wide, sometimes with a narrow wing 2–3 mm wide along the midrib between the primary pinnae, green or greyish, moderately to densely tomentose adaxially and abaxially, the hairs denser abaxially along the midvein, erect, simple, uniseriate, unicellular or multicellular up to 0.3 mm long; margin divided into 4–5 pairs of primary pinnae, the primary pinnae 5–30 x 3–19 mm, decreasing in size from base of leaf to the apex, with (0–) 2–4 irregular secondary pinnae along each side, secondary pinnae (0.5–) 1–5 (–8) x (0.5–) 1–3 (–4.5) mm; the apex of primary and secondary pinnae acute to rounded; petioles 5–32 mm, occasionally with a narrow wing up to 1 mm wide, with indumentum as on the stems.
Inflorescences: 
Inflorescences 4–14 cm long, borne on leafy axillary shoots, leaf opposed, branched 1–2 (–3) times, with 6–22 flowers, all flowers perfect, the axes densely pubescent with simple, uniseriate, multicellular hairs up to 0.4 mm long; peduncle 3.3–5.5 cm long; pedicels 7–10 mm long, not accrescent in fruit, spaced 5–15 mm apart in fruit, articulated at the base.
Flowers: 
Flowers with the calyx 2.9–4.5 (–5.0) mm long, the calyx tube 1.0–2.0 (–2.5) mm long, the lobes 1.0–3.0 (–3.4) x 0.7–1.4 mm at anthesis, 3.0–5.0 x 1.1–2.0 in fruit, elliptic to oblong, glabrous to sparsely pubescent adaxially, moderately to densely pubescent abaxially, especially in the lower half, the hairs simple, uniseriate, unicellular, up to 0.3 mm long, the apex acute to obtuse. Corolla 1.1–2.2 cm in diameter, pentagonal, chartaceous, blue or violet, the tube 1.0–1.5 mm long, the lobes 6–10 x 7–11 mm, sometimes reflexed at anthesis, densely pubescent abaxially with white, simple, uniseriate, unicellular hairs up to 0.2 mm long, glabrous adaxially. Stamens 5, equal; filament tube 1.0–1.5 mm long, glabrous; filaments equal, 0.6–1.0 mm long, glabrous; anthers equal, 3.5–4.6 x 1.0–1.5 mm, oblong to elliptical, yellow, not connivent, poricidal at the tips, the pores tear-drop shaped, tardily opening into longitudinal slits with age. Ovary glabrous; style 8–10 x 0.2–0.3 mm, cylindrical, straight, densely pubescent with white simple, uniseriate, multicellular hairs up to 0.2 mm long, rarely glabrous; stigma clavate, 1.0–1.5 x 0.5–0.8 mm.
Fruits: 
Fruit 4–7 mm in diameter, globose, glabrous, the colour unknown, stone cells absent.
Seeds: 
Seeds 1.5–1.9 x 1.2–1.5 mm, flattened, pale to dark brown, shiny, the surface shallowly pitted.
Chromosome number: 

Not known

Distribution: 

Endemic to Chile [Insula San Ambrosio, Regions II (Antofagasta), III (Atacama) and IV (Coquimbo)]. Arid, rocky or gravelly slopes, quebradas and coastal lomas formations, 20–2200 m in altitude.

Phenology: 
Flowers August to January.
Phylogeny: 

This species belongs to the Regmandra clade of Bohs (2005), an isolated clade within the non-spiny solanums.

Commentary: 

Solanum remyanum is restricted to Chile, and is distinguished by the pinnate, or more usually bipinnate leaves. The pinnae of S. remyanum are broader than those of S. brachyantherum, the other Chilean species with pinnate leaves. Additionally, the flowers are larger than those of S. brachyantherum, in S. remyanum the anthers are ca. 3 mm long compared with ca. 1 mm long in S. brachyantherum. Solanum multifidum from Peru has a similar leaf morphology, but the leaves of S. multifidum are glabrous and the petiole and internodes more prominantly winged; the leaves of S. remyanum are always pubescent and the petioles are usually distinct or with only a narrow wing (up to 1 mm wide).

References: 

Bohs, L. 2005. Major clades in Solanum based on ndhF sequences.
Pp. 27-49 in R. C. Keating, V. C. Hollowell, & T. B. Croat (eds.), A festschrift for William G. D’Arcy: the legacy of a taxonomist. Monographs in Systematic Botany from the Missouri Botanical Garden, Vol. 104. Missouri Botanical Garden Press, St. Louis.

Wed, 2013-11-20 11:02 -- sandy
Scratchpads developed and conceived by (alphabetical): Ed Baker, Katherine Bouton Alice Heaton Dimitris Koureas, Laurence Livermore, Dave Roberts, Simon Rycroft, Ben Scott, Vince Smith